1988- British Society for Developmental Biology (BSDB)

1988- Society for Mathematical Biology (SMB)

1988- Society for Industrial and Applied Mathematics (SIAM)

1991-1996 IMA Specialist Forum on Mathematics in Medicine and Biology

1992- European Society for Mathematical and Theoretical Biology (ESMTB)

1992- London Mathematical Society (LMS)

2000- SIAM Activity Group on Life Sciences

2004-2005 The Physiological Society