Memberships
1988- British Society for Developmental Biology (BSDB)
1988- Society for Mathematical Biology (SMB)
1988- Society for Industrial and Applied Mathematics (SIAM)
1991-1996 IMA Specialist Forum on Mathematics in Medicine and Biology
1992- European Society for Mathematical and Theoretical Biology (ESMTB)
1992- London Mathematical Society (LMS)
2000- SIAM Activity Group on Life Sciences
2004-2005 The Physiological Society